"Which one?" you ask, "and why?" The bottom left registration mark is the problem child. Let me explain.
Sometimes when you are working on a print and cut project you need a little more room in the safe cutting area than what the default registration marks provide. If your design is just slightly outside the safe cutting area you can actually move the registration marks change the margins basically.
Before you move the registration marks though, you should always turn on the 'Show Print Border' option in the Design Page settings. The gray print border line will make it clear what will and what won't print.
When you turn on the Print Border, you can see that the printing area is actually slightly larger than the cut area with the registration makes in their default positions - which is why it's nice you can move them.
Silhouette Studio gives you the option and ability to move the registration marks towards the edge of the page to make more workable room for your design. Watch what happens when I move all of them. The design that was once partially touching the hashed area and was partially outside the cutting area is now completely inside. Wonderful!
BUT there's now a big ol' problem! The problem is...when you move the 'Bottom Inset' registration mark (the bottom left) down - at all - the actual registration mark ends up outside that safe printing area and therefore it will be cut off (aka not print) when your page is printed.
When it's cut off, the Silhouette machine can't read the registration mark or can't read it correctly and therefore you either get a registration failure or a cut that's not accurate.
So the moral of the story is - move the other three registration marks, but hands off the bottom inset or bottom left registration mark.
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